Garen Bohlin has served as a member of our board of directors since July 2010. Since May 2012, Mr. Bohlin has served on the board of directors and as a consultant to multiple life sciences companies. From January 2010 until April 2012, he served as Executive Vice President of Constellation Pharmaceuticals, Inc. Prior to joining Constellation Pharmaceuticals, Mr. Bohlin served as Chief Operating Officer of Sirtris Pharmaceuticals from 2005 to December 2009. Mr. Bohlin was the founding Chief Executive Officer of Syntonix Pharmaceuticals, Inc. from 1999 through December 2005. Earlier in his career, he held multiple executive positions at Genetics Institute and was a partner at Arthur Andersen & Co., a public accounting and consulting organization.. Mr. Bohlin currently serves on the board of directors of Colleqium Pharmaceuticals, Inc., Karyopharm Therapeutics, Inc. and Proteon Therapeutics, Inc.. He also served on the board of directors for Acusphere, Inc. from 2005 to 2014, SpringLeaf Therapeutics, Inc. from 2010 to 2013 and Precision Dermatology, Inc. from 2012-2014. Mr. Bohlin received his B.S. in accounting and finance from The University of Illinois.